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Anhanguera piscator
''Anhanguera piscator'' (meaning "fishing old devil") is a species of pterosaur known from the Early Cretaceous (Aptian age, 112Ma) Santana Formation of Brazil. This pterosaur is closely related to ''Ornithocheirus'', and belongs in the family Ornithocheiridae within its own subfamily, Anhanguerinae.〔Campos, D. de A., and Kellner, A. W. (1985). "Un novo exemplar de ''Anhanguera blittersdorffi'' (Reptilia, Pterosauria) da formaçao Santana, Cretaceo Inferior do Nordeste do Brasil." In Congresso Brasileiro de Paleontologia, Rio de Janeiro, Resumos, p. 13.〕 ''A. piscator'' has also been classified in the genus ''Coloborhynchus'' as ''Coloborhynchus piscator'' or as a synonym of ''Coloborhynchus robustus''.〔Veldmeijer, A.J. (2003). "Description of ''Coloborhynchus spielbergi'' sp. nov. (Pterodactyloidea) from the Albian (Lower Cretaceous) of Brazil." ''Scripta Geologica'', 125: 35-139.〕
==Description==
''Anhanguera piscator'' was a fish-eating animal with a wingspan of about 4.5 m (15 ft). Like many other ornithocheirids, ''A. piscator'' had a rounded crests at front of its upper and lower jaws, which were filled with angled, conical but curved teeth of various sizes and orientations. Like many of its relatives, the jaws were tapered in width, but expanded into a broad, spoon-shaped rosette at the tip. It is distinguished from its relatives by subtle differences in the crest and teeth: unlike its close relatives ''Coloborhynchus'' and ''Ornithocheirus'', the crest on the upper jaw of ''Anhanguera piscator'' did not begin at the tip of the snout, but was set farther back on the skull, and the expanded jaw tips were slender and spoon-shaped rather than robust and box-shaped as in ''Coloborhynchus''. Like many ornithocheiroids, (most notably the pteranodonts but also in ornithocheirids such as ''Ludodactylus'') ''Anhanguera piscator'' had an additional crest protruding from the back of the skull. However, it was reduced to a small, blunt projection in these animals.〔Kellner, A.W.A. and Tomida, Y. (2000). "Description of a new species of Anhangueridae (Pterodactyloidea) with comments on the pterosaurfauna from the Santana Formation (Aptian–Albian), northeastern Brazil." Tokyo, National Science Museum (''National Science Museum Monographs'', 17).〕
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